section_path: Crop alteration and biotechnology/Plant breeding
section_name: Plant breeding
paragraph_text: Plant breeding
  
  Crop alteration has been practiced by humankind for thousands of years, since the beginning of civilization. Altering crops through breeding practices changes the genetic make-up of a plant to develop crops with more beneficial characteristics for humans, for example, larger fruits or seeds, drought-tolerance, or resistance to pests. Significant advances in plant breeding ensued after the work of geneticist Gregor Mendel. His work on dominant and recessive alleles, although initially largely ignored for almost 50 years, gave plant breeders a better understanding of genetics and breeding techniques. Crop breeding includes techniques such as plant selection with desirable traits, self-pollination and cross-pollination, and molecular techniques that genetically modify the organism.
  
  Domestication of plants has, over the centuries increased yield, improved disease resistance and drought tolerance, eased harvest and improved the taste and nutritional value of crop plants. Careful selection and breeding have had enormous effects on the characteristics of crop plants. Plant selection and breeding in the 1920s and 1930s improved pasture (grasses and clover) in New Zealand. Extensive X-ray and ultraviolet induced mutagenesis efforts (i.e. primitive genetic engineering) during the 1950s produced the modern commercial varieties of grains such as wheat, corn (maize) and barley.
  
  The Green Revolution popularized the use of conventional hybridization to sharply increase yield by creating "high-yielding varieties". For example, average yields of corn (maize) in the US have increased from around 2.5 tons per hectare (t/ha) (40 bushels per acre) in 1900 to about 9.4 t/ha (150 bushels per acre) in 2001. Similarly, worldwide average wheat yields have increased from less than 1 t/ha in 1900 to more than 2.5 t/ha in 1990. South American average wheat yields are around 2 t/ha, African under 1 t/ha, and Egypt and Arabia up to 3.5 to 4 t/ha with irrigation. In contrast, the average wheat yield in countries such as France is over 8 t/ha. Variations in yields are due mainly to variation in climate, genetics, and the level of intensive farming techniques (use of fertilizers, chemical pest control, and growth control to avoid lodging).
  
  Investments into innovation for agriculture are long term. This is because it takes time for research to become commercialized and for technology to be adapted to meet multiple regions’ needs, as well as meet national guidelines before being adopted and planted in a farmer’s fields. For instance, it took at least 60 years from the introduction of hybrid corn technology before its adoption became widespread.
  
  Agricultural innovation developed for the specific agroecological conditions of one region is not easily transferred and used in another region with different agroecological conditions. Instead, the innovation would have to be adapted to the specific conditions of that other region and respect its biodiversity and environmental requirements and guidelines. Some such adaptations can be seen through the steadily increasing number of plant varieties protected under the plant variety protection instrument administered by the International Union for the Protection of New Varieties of Plants (UPOV).