section_path: Evolution over geologic timeframes
section_name: Evolution over geologic timeframes
paragraph_text: Evolution over geologic timeframes
  
  Biodiversity is the result of 3.5 billion years of evolution. The origin of life has not been established by science, however, some evidence suggests that life may already have been well-established only a few hundred million years after the formation of the Earth. Until approximately 2.5 billion years ago, all life consisted of microorganisms – archaea, bacteria, and single-celled protozoans and protists.
  
  Biodiversity grew fast during the Phanerozoic (the last 540 million years), especially during the so-called Cambrian explosion —a period during which nearly every phylum of multicellular organisms first appeared. However, recent studies suggest that this diversification had started earlier, at least in the Ediacaran, and that it continued in the Ordovician. Over the next 400 million years or so, invertebrate diversity showed little overall trend and vertebrate diversity shows an overall exponential trend. This dramatic rise in diversity was marked by periodic, massive losses of diversity classified as mass extinction events. A significant loss occurred in anamniotic limbed vertebrates when rainforests collapsed in the Carboniferous, but amniotes seem to have been little affected by this event; their diversification slowed down later, around the Asselian / Sakmarian boundary, in the early Cisuralian (Early Permian), about 293 Ma ago. The worst was the Permian-Triassic extinction event, 251 million years ago. Vertebrates took 30 million years to recover from this event.
  
  The most recent major mass extinction event, the Cretaceous–Paleogene extinction event, occurred 66 million years ago. This period has attracted more attention than others because it resulted in the extinction of the non-avian dinosaurs, which were represented by many lineages at the end of the Maastrichtian, just before that extinction event. However, many other taxa were affected by this crisis, which affected even marine taxa, such as ammonites, which also became extinct around that time.
  
  The biodiversity of the past is called Paleobiodiversity. The fossil record suggests that the last few million years featured the greatest biodiversity in history. However, not all scientists support this view, since there is uncertainty as to how strongly the fossil record is biased by the greater availability and preservation of recent geologic sections. Some scientists believe that corrected for sampling artifacts, modern biodiversity may not be much different from biodiversity 300 million years ago, whereas others consider the fossil record reasonably reflective of the diversification of life. Estimates of the present global macroscopic species diversity vary from 2 million to 100 million, with a best estimate of somewhere near 9 million, the vast majority arthropods. Diversity appears to increase continually in the absence of natural selection.