section_path: Ecology
section_name: Ecology
paragraph_text: Ecology
  
  Forests account for 75% of the gross primary productivity of the Earth's biosphere, and contain 80% of the Earth's plant biomass. Biomass per unit area is high compared to other vegetation communities. Much of this biomass occurs below ground in the root systems and as partially decomposed plant detritus. The woody component of a forest contains lignin, which is relatively slow to decompose compared with other organic materials such as cellulose or carbohydrate. The world's forests contain about 606 gigatonnes of living biomass (above- and below-ground) and 59 gigatonnes of dead wood. The total biomass has decreased slightly since 1990, but biomass per unit area has increased.
  
  Forest ecosystems broadly differ based on climate; latitudes 10° north and south of the equator are mostly covered in tropical rainforest, and the latitudes between 53°N and 67°N have boreal forest. As a general rule, forests dominated by angiosperms (broadleaf forests) are more species-rich than those dominated by gymnosperms (conifer, montane, or needleleaf forests), although exceptions exist. The trees that form the principal structural and defining component of a forest may be of a great variety of species (as in tropical rainforests and temperate deciduous forests), or relatively few species over large areas (e.g., taiga and arid montane coniferous forests). The biodiversity of forests also encompasses shrubs, herbaceous plants, mosses, ferns, lichens, fungi, and a variety of animals.
  
  Trees rising up to 35 meters (115 ft) in height add a vertical dimension to the area of land that can support plant and animal species, opening up numerous ecological niches for arboreal animal species, epiphytes, and various species that thrive under the regulated microclimate created under the canopy. Forests have intricate three-dimensional structures that increase in complexity with lower levels of disturbance and greater variety of tree species.
  
  The biodiversity of forests varies considerably according to factors such as forest type, geography, climate, and soils – in addition to human use. Most forest habitats in temperate regions support relatively few animal and plant species, and species that tend to have large geographical distributions, while the montane forests of Africa, South America, Southeast Asia, and lowland forests of Australia, coastal Brazil, the Caribbean islands, Central America, and insular Southeast Asia have many species with small geographical distributions. Areas with dense human populations and intense agricultural land use, such as Europe, parts of Bangladesh, China, India, and North America, are less intact in terms of their biodiversity. Northern Africa, southern Australia, coastal Brazil, Madagascar, and South Africa are also identified as areas with striking losses in biodiversity intactness.