section_path: History/Early history
section_name: Early history
paragraph_text: Early history
  
  The first human settlement on Russia dates back to the Oldowan period in the early Lower Paleolithic. About 2 million years ago, representatives of Homo erectus migrated to the Taman Peninsula in southern Russia. Flint tools, some 1.5 million years old, have been discovered in the North Caucasus. Radiocarbon dated specimens from Denisova Cave in the Altai Mountains estimate the oldest Denisovan specimen lived 195–122,700 years ago. Fossils of Denny, an archaic human hybrid that was half Neanderthal and half Denisovan, and lived some 90,000 years ago, was also found within the latter cave. Russia was home to some of the last surviving Neanderthals, from about 45,000 years ago, found in Mezmaiskaya cave.
  
  The first trace of an early modern human in Russia dates back to 45,000 years, in Western Siberia. The discovery of high concentration cultural remains of anatomically modern humans, from at least 40,000 years ago, was found at Kostyonki–Borshchyovo, and at Sungir, dating back to 34,600 years ago—both in western Russia. Humans reached Arctic Russia at least 40,000 years ago, in Mamontovaya Kurya. Ancient North Eurasian populations from Siberia genetically similar to Mal'ta–Buret' culture and Afontova Gora were an important genetic contributor to Ancient Native Americans and Eastern Hunter-Gatherers.
  
  The Kurgan hypothesis places the Volga-Dnieper region of southern Russia and Ukraine as the urheimat of the Proto-Indo-Europeans. Early Indo-European migrations from the Pontic–Caspian steppe of Ukraine and Russia spread Yamnaya ancestry and Indo-European languages across large parts of Eurasia. Nomadic pastoralism developed in the Pontic–Caspian steppe beginning in the Chalcolithic. Remnants of these steppe civilisations were discovered in places such as Ipatovo, Sintashta, Arkaim, and Pazyryk, which bear the earliest known traces of horses in warfare. The genetic makeup of speakers of the Uralic language family in northern Europe was shaped by migration from Siberia that began at least 3,500 years ago.
  
  In the 3rd to 4th centuries CE, the Gothic kingdom of Oium existed in southern Russia, which was later overrun by Huns. Between the 3rd and 6th centuries CE, the Bosporan Kingdom, which was a Hellenistic polity that succeeded the Greek colonies, was also overwhelmed by nomadic invasions led by warlike tribes such as the Huns and Eurasian Avars. The Khazars, who were of Turkic origin, ruled the steppes between the Caucasus in the south, to the east past the Volga river basin, and west as far as Kyiv on the Dnieper river until the 10th century. After them came the Pechenegs who created a large confederacy, which was subsequently taken over by the Cumans and the Kipchaks.
  
  The ancestors of Russians are among the Slavic tribes that separated from the Proto-Indo-Europeans, who appeared in the northeastern part of Europe c. 1500 years ago. The East Slavs gradually settled western Russia (approximately between modern Moscow and Saint-Petersburg) in two waves: one moving from Kiev towards present-day Suzdal and Murom and another from Polotsk towards Novgorod and Rostov. Prior to Slavic migration, that territory was populated by Finno-Ugrian peoples. From the 7th century onwards, the incoming East Slavs slowly assimilated the native Finno-Ugrians.