History.

The Kingdom of Asturias began in 720, with the Visigothic aristocrat Pelagius's (685–737) revolt against the Muslims who at the time were occupying most of the Iberian Peninsula. The Moorish invasion that began in 711 had taken control of most of the peninsula, until the revolt in the northern mountains by Pelagius. The resulting Kingdom of Asturias, located in an economically poor region of Iberia, was largely ignored by the Muslims. In 720, the area where Oviedo is now located was still uninhabited.  It is said that two monks, Máximo and Fromestano (), founded the city in 761. That settlement was soon to be completed with the construction of a small church dedicated to Saint Vincent. Oviedo was established on an uninhabited hillside, with no Visigothic or Roman foundation before it became an Asturian city. Following Pelagius, who died in 737, Alfonso I (739–57) founded a dynasty that would last until 1037. The Asturian Kingdom was on hostile terms with southern Moorish Spain. In 794, Oviedo was sacked and pillaged by Caliph Hisham I in one of his numerous campaigns against the Christian kingdoms.  King Alfonso I is said to have "set in place the whole order of the Goths, as it had been in Toledo, as much in the church as in the palace." The intention with Oviedo was to shape it into a city similar to that of Visigothic Toledo. Once kings had settled in Oviedo, they adopted as much of the architectural style and imagery of Toledo. Even with this in mind, Oviedo did not necessarily resemble the old Visigothic capital in Toledo. The churches and buildings of Oviedo follow instead late provincial Roman tradition. Since Asturias at the time was an agriculturally poor area of Spain the scale of the buildings is quite impressive.  Oviedo's rich architectural tradition began with King Fruela I (757–768). King Fruela I of Asturias, the fourth of the Asturian monarchs, was the first decided promoter of the city as may be witnessed by his construction of both a palace and a nearby church. This church was later restored by Alfonso II. Oviedo owes to a later king, Alfonso II The Chaste (791–842), its establishment as a capital city and ruling seat as a result of the moving of the court from Pravia and the creation of the Pilgrim's Route to Santiago de Compostela, a major event in the history of Oviedo, a church dedicated to The Saviour, the Cathedral of San Salvador, and a royal palace formed the nucleus of Oviedo. Also constructed during Alfonso II's reign was the San Julian de los Prados church, which is one of the best preserved Asturian churches. Alfonso II's successor, Ramiro I (842–850), continued Alfonso II's construction streak. Ramiro I constructed two buildings, the Church Santa Maria del Naranco and San Miguel de Lillo. The Church Santa Maria de Naranco was likely to originally be Ramiro I's palace and later changed into a church. By this time the Court of the Palace was centered in Oviedo, which was the main royal residence. This court was controlled by member of the Asturian nobility.  Ramiro I's (842–850) eight-year tenure was uneasy, he faced rebellions from the Counts of the Palace. The first rebellion against Ramiro I was led by Alroitus, and the second rebellion was led by Piniolus. Both of these rebellions were unsuccessful in removing Ramiro I. These rebellions may have been why Ramiro I built his palace in the mountains surrounding Oviedo, presumably away from the violence. During the 9th century in Oviedo, Roman style property law is common. 9th century documents also indicate small scale aristocracies across the kingdom, as well as a large presence of a landowning peasantry.  Following Ramiro I's reign, Ordoño I (850–866) came into power and began the Asturian king's father-son succession. Ordoño I was the first king to push southwards into Arab territory. Following Ordoño I's death on May 27, 866, usurpers attempted to take the throne. The following king Alfonso III (866–910), who was thirteen at the time, took refuge in Castile until his followers had killed the usurper.  Alfonso III's contributions to building construction are not nearly as well documented as Ramiro I's or Alfonso II's contributions. The Chronicle of Alfonso III does not mention any buildings created by Alfonso III, neither does the Chronicle of Albelda. In 882, the body of the Cordoban martyr Eulogius was sent to Oviedo. This was meant a diplomatic gift from Emir Muhammad I (852–886). Eulogius was executed in 859. The body was likely accompanied by Eulogius's book collection. In the 16th century, the only manuscript of Eulogius's writings was discovered in the Oviedo Cathedral Library. Here it was copied once before it disappeared completely from the library. Following an offensive in 881 against an Umayyad army, Alfsonso III returned to Oviedo to rebuild churches. It was at this time he constructed one or more palaces. The Chronicle of Albelda and the Chronicle of Sampiro tie Alfonso III's victories in battle to his program of church building in Oviedo. In 908, Alfonso III commissioned a gold and jewelled cross to contain the cross carried by Pelagius I at Covadonga. This "Cross of Victory" is located in the Camara Sancta in the Oviedo Cathedral. However, recent Carbon14 analysis of the wooden cross indicates that it was no older than the golden casing created to surround the cross. The commission of the casing shows us Alfonso III's interest in perpetuating the legend of Pelagius I.  Towards the end of Alfonso III's reign, he faced many challenges. In 901, a prophet named Alhaman led a "great army of Muslims" and attempted to take Zamora. To add to this, Alfonso III's brother Vermudo revolted in Astorga. There were several attempts at the aging Alfonso III's life by his sons. Alfonso III was overthrown by sons, and died in Zamora. His body was taken to Oviedo for burial.  The moving of the royal court to León, after the death of Alfonso III, 'The Great', links the life of the city to the relics preserved in its cathedral and the passing of pilgrims that visit El Salvador, and continue on their way to Santiago de Compostela. Kings spent less and less time in Oviedo following the change, and spent more time in the rich Duero Plains. León was built up after it became the capital, and eventually surpassed Oviedo in terms of construction.  During the 12th century, many Royal Charters were fabricated by Bishop Pelayo de Oviedo, "el fabulador" ("the fabulist"). Since were few checks on internal bookkeeping in the Asturian kingdom actions like this were commonplace in the kingdom. When original documents faded, they were copied onto cartularies and often with alterations that suited the needs of those who copied the documents. The most glaring example can be seen in the "Liber Testamentorum", which was compiled by Bishop Pelayo de Oviedo in 1109. This document contained many confirmation rights and property rights of the Oviedo cathedral by Asturian and Leonese Kings. Bishop Pelayo's intent behind this was to try to gain the independence of his see from the archbishop of Toledo or Santiago, as well as to promote Oviedo as a pilgrim destination. According to Sánchez-Albornoz, "He (Bishop Pelayo) always, always, always falsified." It is assumed that Bishop Pelayo never committed forgery for the enjoyment, but primarily to promote the church of Oviedo.  The following centuries (12th–16th) witness the development of the medieval city, the outlines of which are still preserved today, the construction of the city walls, a devastating fire which took place on Christmas Eve in 1521, and the aqueduct works, "Los Pilares", constructed in order to provide the city with water throughout the 16th century.  The foundation of the Arts College (University of Oviedo) by Fernando de Valdés Salas, at the beginning of the 17th century, opened Oviedo to a progressive urban expansion. Further impulse was in the 18th century by the regional nobility and the construction of remarkable palaces; in the 19th century by industrial growth and the suburban development of Uría Street; and finally in the 20th century by administrative and commercial development.  In October 1934, there was a left-wing revolt against the conservative government, based in several cities. In Asturias, the fighting developed into a small, short-lived civil war: the Asturian miners' strike of 1934. 50,000 workers, mostly miners, armed themselves with dynamite and captured Oviedo after heavy fighting. They gained control of the arsenal with 30,000 rifles and machine guns. The Army Chief of Staff, General Francisco Franco sent in soldiers who overpowered the rebels after severe street fighting that left 3,000 rebels dead and 7,000 wounded. The cathedral was badly damaged, with its eighth-century chapel blown up by a mine. In the aftermath, many false atrocity stories circulated.  The Siege of Oviedo in 1936 was a memorable event in the Spanish Civil War. The army garrison rose in support of the Nationalist coup d'état and withstood a siege of three months by an improvised Republican force until relieved in 1937.

Geography.

Oviedo is located in the centre of Asturias between the Nalón River and Nora River. To the north lie Las Regueras and Llanera, to the south Mieres and Ribera de Arriba, to the east Siero and Langreo, and to the west Grado and Santo Adriano. The altitude of Oviedo is between 80 and 709 metres above sea level. The city is protected against strong winds by Monte Naranco in the north and the Sierra del Aramo in the south. The city centre is rather hilly.

Economy.

The economy is strongly dependent on the service sector, with many office buildings in the city centre. Oviedo's status as the administrative centre of the region supports a large number of jobs in public administration. The manufacturing sector, which remains important in this part of Spain, is not prevalent in Oviedo itself, but is more important in the adjacent municipalities of Siero and Llanera which lie to the north of the city, between Oviedo and Gijon. In 2009, the Oviedo municipality had a total debt of €135,185,000.00.

Architecture.

Oviedo contains a very rich architectural history, with many buildings dating back to the early medieval period. Many of the building projects were undertaken during Alfonso II's (791-842) reign and Ramiro I's (842-850) reign. Alfonso III's contributions are not as well documented.  Alfonso II is said to have built four churches, one dedicated to Christ the Saviour, the Blessed Virgin Mary, St. Tyrsus, and SS Julian and Basilissa. There are few traces of the churches dedicated to the Saviour, the Virgin Mary, and St. Tyrsus. The San Salvadore church, which was dedicated to the Saviour, is likely beneath the Cathedral of Oviedo. The church of Santa Maria de la Corte, which was dedicated to the Virgin Mary, was demolished in 1702. As for St. Tyrsus, the church dedicated to him exists today as the church of San Tirso. Only a wall and a three light window are believed to have been built by Alfonso II, the majority of the rest of the church is dated to the 14th century. The best preserved church constructed during Alfonso II's time was San Julian de los Prados.  Two buildings are said to have been built during Ramiro I's reign, one was a church dedicated to the Virgin Mary and the other construction was a palace. These were built just outside Oviedo, on Monte Lignum. The church Santa Maria de Naranco seems to originally have been a palace, but later repurposed into a church. The church has an atypical plan from other churches at its time, possibly because it was supposed to contain a throne room for the king. The other church built during Ramiro I's time was San Miguel de Lillo.  The Chronicle of Albelda, one of the primary sources used to discern which King commissioned which building, only extends to 883. Because of this, constructions undertaken during Alfonso III's time as king were not documented.  The following is a list of the notable architectural sites in Oviedo:

Culture.

Oviedo inspired the fictional city of "Vetusta" in Leopoldo Alas's "La Regenta". Other Spanish writers were inspired by the city, including Ramón Pérez de Ayala in "Tigre Juan" and Dolores Medio in her novel "Nosotros los Rivero".  Oviedo was featured prominently in Woody Allen's movie "Vicky Cristina Barcelona".  Museums in Oviedo:  Other cultural centres in Oviedo:  Cultural institutions: Orquesta Sinfonica del Principado de Asturias is the premier orchestra of the region, the Principality of Asturias. This full-time symphony orchestra performs a wide range of the classical repertoire with world class soloists and conductors. It is based in the Auditorio Principe Felipe in Oviedo, but it also performs regularly at the main concert venues in Gijón and Avilés. It is Internationally recognized as one of the best orchestras in Spain, it is also committed to adventurous programing with strong emphasis on education and community partnerships. Rossen Milanov is the Music Director.  Oviedo also hosts the annual Princess of Asturias Awards (previously called the Prince of Asturias Awards). This prestigious event, held in the city's Campoamor Theatre, recognizes international achievement in eight different categories. Previous award winners include Oscar Niemeyer, Bob Dylan and Francis Ford Coppola in the category of Arts; Nelson Mandela, the International Space Station and Al Gore in the category of International Cooperation; and Mario Bunge, CNN and Quino in the category of Communications and Humanities.  Oviedo University's international campus attracts many foreign scholars from all over the globe.  The city lends its name to the sudarium of Oviedo a religious relic revered there since the 9th century.

Politics.

The current mayor is Alfredo Canteli (PP).

Sports.

Formula One.

The most famous athlete from Oviedo is 2005 and 2006 Formula One world champion Fernando Alonso, famous for being Spain's only Formula One title winner, having raced for teams like Renault, Ferrari and McLaren. Alonso has his official career and life museum, together with a karting circuit designed by himself, in Llanera, nearby Oviedo.

Feasts and traditions.

The Festive Calendar:  Typical gastronomy of the province:

Climate.

Oviedo's climate is temperate oceanic (Cfb in the Köppen climate classification). Its climate is very similar to neighbouring city Gijón, with only narrow fluctuations in temperature. Oviedo's hottest month is in August with an average high of. The city centre is located at a lower elevation than the weather station so is likely somewhat milder year-round. Its maritime position renders winters much milder than in continental Spain such as in the Madrid capital region, but summers naturally are far less hot than in the interior. There is a slight drying tendency during summer, albeit far less significant than in other areas of Spain.

Transport.

Rail.

The Oviedo railway station provides a wide range of long- and middle-distance services, in addition to regional and suburban ("cercanías") services operated by Renfe, and the narrow-gauge Renfe Feve lines. Within the municipality, there are others rail stations on the Cercanías Asturias network, in La Corredoria, Llamaquique, El Caleyo, Olloniego, Santa Eulalia de Manzaneda, Tudela-Veguín, Parque Principado, Colloto, Argañosa-Lavapiés, Las Campas, Las Mazas, San Claudio, Soto-Udrión, Trubia, Caces, Fuso de la Reina, and Caces.

Tramway.

From 1922 to 1956, there was a tramway system in Oviedo

International relations.

Twin towns — sister cities.

Oviedo is twinned with:

Oviedo (,, ; ) is the capital city of the Principality of Asturias in northern Spain and the administrative and commercial centre of the region. It is also the name of the municipality that contains the city. Oviedo is located approximately southwest of Gijón and south of Avilés, both of which lie on the shoreline of the Bay of Biscay. Its proximity to the ocean causes Oviedo to have a maritime climate, in spite of its not being located on the shoreline itself.

Oviedo (asturisch: "Uviéu" oder "Uvieo") ist die Hauptstadt der Autonomen Gemeinschaft Fürstentum Asturien im Norden Spaniens. Im 9. und 10. Jahrhundert war die Stadt die Residenz des damals unabhängigen Königreiches Asturien. Die Bewohner der Stadt werden „Ovetenser“ genannt.


Aims.

The field of plant physiology includes the study of all the internal activities of plants—those chemical and physical processes associated with life as they occur in plants. This includes study at many levels of scale of size and time. At the smallest scale are molecular interactions of photosynthesis and internal diffusion of water, minerals, and nutrients. At the largest scale are the processes of plant development, seasonality, dormancy, and reproductive control. Major subdisciplines of plant physiology include phytochemistry (the study of the biochemistry of plants) and phytopathology (the study of disease in plants). The scope of plant physiology as a discipline may be divided into several major areas of research. First, the study of phytochemistry (plant chemistry) is included within the domain of plant physiology. To function and survive, plants produce a wide array of chemical compounds not found in other organisms. Photosynthesis requires a large array of pigments, enzymes, and other compounds to function. Because they cannot move, plants must also defend themselves chemically from herbivores, pathogens and competition from other plants. They do this by producing toxins and foul-tasting or smelling chemicals. Other compounds defend plants against disease, permit survival during drought, and prepare plants for dormancy, while other compounds are used to attract pollinators or herbivores to spread ripe seeds.  Secondly, plant physiology includes the study of biological and chemical processes of individual plant cells. Plant cells have a number of features that distinguish them from cells of animals, and which lead to major differences in the way that plant life behaves and responds differently from animal life. For example, plant cells have a cell wall which restricts the shape of plant cells and thereby limits the flexibility and mobility of plants. Plant cells also contain chlorophyll, a chemical compound that interacts with light in a way that enables plants to manufacture their own nutrients rather than consuming other living things as animals do.  Thirdly, plant physiology deals with interactions between cells, tissues, and organs within a plant. Different cells and tissues are physically and chemically specialized to perform different functions. Roots and rhizoids function to anchor the plant and acquire minerals in the soil. Leaves catch light in order to manufacture nutrients. For both of these organs to remain living, minerals that the roots acquire must be transported to the leaves, and the nutrients manufactured in the leaves must be transported to the roots. Plants have developed a number of ways to achieve this transport, such as vascular tissue, and the functioning of the various modes of transport is studied by plant physiologists.  Fourthly, plant physiologists study the ways that plants control or regulate internal functions. Like animals, plants produce chemicals called hormones which are produced in one part of the plant to signal cells in another part of the plant to respond. Many flowering plants bloom at the appropriate time because of light-sensitive compounds that respond to the length of the night, a phenomenon known as photoperiodism. The ripening of fruit and loss of leaves in the winter are controlled in part by the production of the gas Ethylene as a plant hormone#ethylene by the plant.  Finally, plant physiology includes the study of plant response to environmental conditions and their variation, a field known as environmental physiology. Stress from water loss, changes in air chemistry, or crowding by other plants can lead to changes in the way a plant functions. These changes may be affected by genetic, chemical, and physical factors.

Biochemistry of plants.

The chemical elements of which plants are constructed—principally carbon, oxygen, hydrogen, nitrogen, phosphorus, sulfur, etc.—are the same as for all other life forms animals, fungi, bacteria and even viruses. Only the details of the molecules into which they are assembled differs.  Despite this underlying similarity, plants produce a vast array of chemical compounds with unique properties which they use to cope with their environment. Pigments are used by plants to absorb or detect light, and are extracted by humans for use in dyes. Other plant products may be used for the manufacture of commercially important rubber or biofuel. Perhaps the most celebrated compounds from plants are those with pharmacological activity, such as salicylic acid from which aspirin is made, morphine, and digoxin. Drug companies spend billions of dollars each year researching plant compounds for potential medicinal benefits.

Constituent elements.

Plants require some nutrients, such as carbon and nitrogen, in large quantities to survive. Some nutrients are termed macronutrients, where the prefix "macro-" (large) refers to the quantity needed, not the size of the nutrient particles themselves. Other nutrients, called micronutrients, are required only in trace amounts for plants to remain healthy. Such micronutrients are usually absorbed as ions dissolved in water taken from the soil, though carnivorous plants acquire some of their micronutrients from captured prey.  The following tables list element nutrients essential to plants. Uses within plants are generalized.

Pigments.

Among the most important molecules for plant function are the pigments. Plant pigments include a variety of different kinds of molecules, including porphyrins, carotenoids, and anthocyanins. All biological pigments selectively absorb certain wavelengths of light while reflecting others. The light that is absorbed may be used by the plant to power chemical reactions, while the reflected wavelengths of light determine the color the pigment appears to the eye.  Chlorophyll is the primary pigment in plants; it is a porphyrin that absorbs red and blue wavelengths of light while reflecting green. It is the presence and relative abundance of chlorophyll that gives plants their green color. All land plants and green algae possess two forms of this pigment: chlorophyll "a" and chlorophyll "b". Kelps, diatoms, and other photosynthetic heterokonts contain chlorophyll "c" instead of "b", red algae possess chlorophyll "a". All chlorophylls serve as the primary means plants use to intercept light to fuel photosynthesis.  Carotenoids are red, orange, or yellow tetraterpenoids. They function as accessory pigments in plants, helping to fuel photosynthesis by gathering wavelengths of light not readily absorbed by chlorophyll. The most familiar carotenoids are carotene (an orange pigment found in carrots), lutein (a yellow pigment found in fruits and vegetables), and lycopene (the red pigment responsible for the color of tomatoes). Carotenoids have been shown to act as antioxidants and to promote healthy eyesight in humans.  Anthocyanins (literally "flower blue") are water-soluble flavonoid pigments that appear red to blue, according to pH. They occur in all tissues of higher plants, providing color in leaves, stems, roots, flowers, and fruits, though not always in sufficient quantities to be noticeable. Anthocyanins are most visible in the petals of flowers, where they may make up as much as 30% of the dry weight of the tissue. They are also responsible for the purple color seen on the underside of tropical shade plants such as "Tradescantia zebrina". In these plants, the anthocyanin catches light that has passed through the leaf and reflects it back towards regions bearing chlorophyll, in order to maximize the use of available light  Betalains are red or yellow pigments. Like anthocyanins they are water-soluble, but unlike anthocyanins they are indole-derived compounds synthesized from tyrosine. This class of pigments is found only in the Caryophyllales (including cactus and amaranth), and never co-occur in plants with anthocyanins. Betalains are responsible for the deep red color of beets, and are used commercially as food-coloring agents. Plant physiologists are uncertain of the function that betalains have in plants which possess them, but there is some preliminary evidence that they may have fungicidal properties.

Signals and regulators.

Plants produce hormones and other growth regulators which act to signal a physiological response in their tissues. They also produce compounds such as phytochrome that are sensitive to light and which serve to trigger growth or development in response to environmental signals.

Plant hormones.

Plant hormones, known as plant growth regulators (PGRs) or phytohormones, are chemicals that regulate a plant's growth. According to a standard animal definition, hormones are signal molecules produced at specific locations, that occur in very low concentrations, and cause altered processes in target cells at other locations. Unlike animals, plants lack specific hormone-producing tissues or organs. Plant hormones are often not transported to other parts of the plant and production is not limited to specific locations.  Plant hormones are chemicals that in small amounts promote and influence the growth, development and differentiation of cells and tissues. Hormones are vital to plant growth; affecting processes in plants from flowering to seed development, dormancy, and germination. They regulate which tissues grow upwards and which grow downwards, leaf formation and stem growth, fruit development and ripening, as well as leaf abscission and even plant death.  The most important plant hormones are abscissic acid (ABA), auxins, ethylene, gibberellins, and cytokinins, though there are many other substances that serve to regulate plant physiology.

Photomorphogenesis.

While most people know that light is important for photosynthesis in plants, few realize that plant sensitivity to light plays a role in the control of plant structural development (morphogenesis). The use of light to control structural development is called photomorphogenesis, and is dependent upon the presence of specialized photoreceptors, which are chemical pigments capable of absorbing specific wavelengths of light.  Plants use four kinds of photoreceptors: phytochrome, cryptochrome, a UV-B photoreceptor, and protochlorophyllide "a". The first two of these, phytochrome and cryptochrome, are photoreceptor proteins, complex molecular structures formed by joining a protein with a light-sensitive pigment. Cryptochrome is also known as the UV-A photoreceptor, because it absorbs ultraviolet light in the long wave "A" region. The UV-B receptor is one or more compounds not yet identified with certainty, though some evidence suggests carotene or riboflavin as candidates. Protochlorophyllide "a", as its name suggests, is a chemical precursor of chlorophyll.  The most studied of the photoreceptors in plants is phytochrome. It is sensitive to light in the red and far-red region of the visible spectrum. Many flowering plants use it to regulate the time of flowering based on the length of day and night (photoperiodism) and to set circadian rhythms. It also regulates other responses including the germination of seeds, elongation of seedlings, the size, shape and number of leaves, the synthesis of chlorophyll, and the straightening of the epicotyl or hypocotyl hook of dicot seedlings.

Photoperiodism.

Many flowering plants use the pigment phytochrome to sense seasonal changes in day length, which they take as signals to flower. This sensitivity to day length is termed photoperiodism. Broadly speaking, flowering plants can be classified as long day plants, short day plants, or day neutral plants, depending on their particular response to changes in day length. Long day plants require a certain minimum length of daylight to starts flowering, so these plants flower in the spring or summer. Conversely, short day plants flower when the length of daylight falls below a certain critical level. Day neutral plants do not initiate flowering based on photoperiodism, though some may use temperature sensitivity (vernalization) instead.  Although a short day plant cannot flower during the long days of summer, it is not actually the period of light exposure that limits flowering. Rather, a short day plant requires a minimal length of uninterrupted darkness in each 24-hour period (a short daylength) before floral development can begin. It has been determined experimentally that a short day plant (long night) does not flower if a flash of phytochrome activating light is used on the plant during the night.  Plants make use of the phytochrome system to sense day length or photoperiod. This fact is utilized by florists and greenhouse gardeners to control and even induce flowering out of season, such as the "Poinsettia".

Environmental physiology.

Paradoxically, the subdiscipline of environmental physiology is on the one hand a recent field of study in plant ecology and on the other hand one of the oldest. Environmental physiology is the preferred name of the subdiscipline among plant physiologists, but it goes by a number of other names in the applied sciences. It is roughly synonymous with ecophysiology, crop ecology, horticulture and agronomy. The particular name applied to the subdiscipline is specific to the viewpoint and goals of research. Whatever name is applied, it deals with the ways in which plants respond to their environment and so overlaps with the field of ecology.  Environmental physiologists examine plant response to physical factors such as radiation (including light and ultraviolet radiation), temperature, fire, and wind. Of particular importance are water relations (which can be measured with the Pressure bomb) and the stress of drought or inundation, exchange of gases with the atmosphere, as well as the cycling of nutrients such as nitrogen and carbon.  Environmental physiologists also examine plant response to biological factors. This includes not only negative interactions, such as competition, herbivory, disease and parasitism, but also positive interactions, such as mutualism and pollination.

Tropisms and nastic movements.

Plants may respond both to directional and non-directional stimuli. A response to a directional stimulus, such as gravity or sunlight, is called a tropism. A response to a nondirectional stimulus, such as temperature or humidity, is a nastic movement.  Tropisms in plants are the result of differential cell growth, in which the cells on one side of the plant elongates more than those on the other side, causing the part to bend toward the side with less growth. Among the common tropisms seen in plants is phototropism, the bending of the plant toward a source of light. Phototropism allows the plant to maximize light exposure in plants which require additional light for photosynthesis, or to minimize it in plants subjected to intense light and heat. Geotropism allows the roots of a plant to determine the direction of gravity and grow downwards. Tropisms generally result from an interaction between the environment and production of one or more plant hormones.  Nastic movements results from differential cell growth (e.g. epinasty and hiponasty), or from changes in turgor pressure within plant tissues (e.g., nyctinasty), which may occur rapidly. A familiar example is thigmonasty (response to touch) in the Venus fly trap, a carnivorous plant. The traps consist of modified leaf blades which bear sensitive trigger hairs. When the hairs are touched by an insect or other animal, the leaf folds shut. This mechanism allows the plant to trap and digest small insects for additional nutrients. Although the trap is rapidly shut by changes in internal cell pressures, the leaf must grow slowly to reset for a second opportunity to trap insects.

Plant disease.

Economically, one of the most important areas of research in environmental physiology is that of phytopathology, the study of diseases in plants and the manner in which plants resist or cope with infection. Plant are susceptible to the same kinds of disease organisms as animals, including viruses, bacteria, and fungi, as well as physical invasion by insects and roundworms.  Because the biology of plants differs with animals, their symptoms and responses are quite different. In some cases, a plant can simply shed infected leaves or flowers to prevent the spread of disease, in a process called abscission. Most animals do not have this option as a means of controlling disease. Plant diseases organisms themselves also differ from those causing disease in animals because plants cannot usually spread infection through casual physical contact. Plant pathogens tend to spread via spores or are carried by animal vectors.  One of the most important advances in the control of plant disease was the discovery of Bordeaux mixture in the nineteenth century. The mixture is the first known fungicide and is a combination of copper sulfate and lime. Application of the mixture served to inhibit the growth of downy mildew that threatened to seriously damage the French wine industry.

History.

Early history.

[Francis Bacon]] published one of the first plant physiology experiments in 1627 in the book, "Sylva Sylvarum." Bacon grew several terrestrial plants, including a rose, in water and concluded that soil was only needed to keep the plant upright. Jan Baptist van Helmont published what is considered the first quantitative experiment in plant physiology in 1648. He grew a willow tree for five years in a pot containing 200 pounds of oven-dry soil. The soil lost just two ounces of dry weight and van Helmont concluded that plants get all their weight from water, not soil. In 1699, John Woodward published experiments on growth of spearmint in different sources of water. He found that plants grew much better in water with soil added than in distilled water.  Stephen Hales is considered the Father of Plant Physiology for the many experiments in the 1727 book,"Vegetable Staticks"; though Julius von Sachs unified the pieces of plant physiology and put them together as a discipline. His "Lehrbuch der Botanik" was the plant physiology bible of its time.  Researchers discovered in the 1800s that plants absorb essential mineral nutrients as inorganic ions in water. In natural conditions, soil acts as a mineral nutrient reservoir but the soil itself is not essential to plant growth. When the mineral nutrients in the soil are dissolved in water, plant roots absorb nutrients readily, soil is no longer required for the plant to thrive. This observation is the basis for hydroponics, the growing of plants in a water solution rather than soil, which has become a standard technique in biological research, teaching lab exercises, crop production and as a hobby.

Economic applications.

Food production.

In horticulture and agriculture along with food science, plant physiology is an important topic relating to fruits, vegetables, and other consumable parts of plants. Topics studied include: "climatic" requirements, fruit drop, nutrition, ripening, fruit set. The production of food crops also hinges on the study of plant physiology covering such topics as optimal planting and harvesting times and post harvest storage of plant products for human consumption and the production of secondary products like drugs and cosmetics.

Plant physiology is a subdiscipline of botany concerned with the functioning, or physiology, of plants. Closely related fields include plant morphology (structure of plants), plant ecology (interactions with the environment), phytochemistry (biochemistry of plants), cell biology, genetics, biophysics and molecular biology.

Die Pflanzenphysiologie ist die Wissenschaft von den Lebensvorgängen (Physiologie) der Pflanzen. Ein zentraler Vorgang ist die Photosynthese, an die sich die Bildung anderer Substanzen (von der Glucose über Polysaccharide bis zu Lipiden, Proteinen und Nukleinsäuren) anschließt. Außerdem werden Wachstumsprozesse, die Differenzierung von Organen, Reaktionen auf Umweltreize, Stofftransporte und die Kommunikation zwischen Zellen, Geweben und Organen untersucht.


History.

Early history.

Plains in Hebei were the home of Peking man, a group of Homo erectus that lived in the area around 200,000 to 700,000 years ago. Neolithic findings at the prehistoric Beifudi site date back to 7000 and 8000 BC.

Pre-Qin dynasty era.

During the Spring and Autumn period (722 BC – 476 BC), Hebei was under the rule of the states of Yan in the north and Jin in the south. Also during this period, a nomadic people known as Dí invaded the plains of northern China and established Zhongshan in central Hebei. During the Warring States period (403 BC–221 BC), Jin was partitioned, and much of its territory within Hebei went to Zhao.

Qin and Han dynasties.

The Qin dynasty unified China in 221 BC. The Han dynasty (206 BC – AD 220) ruled the area under two provinces ("zhou"), You Prefecture in the north and Ji Province in the south. At the end of the Han dynasty, most of Hebei came under the control of warlords Gongsun Zan in the north and Yuan Shao further south; Yuan Shao emerged victorious of the two, but he was soon defeated by rival Cao Cao (based further south, in modern-day Henan) in the Battle of Guandu in 200. Hebei then came under the rule of the Kingdom of Wei (one of the Three Kingdoms), established by the descendants of Cao Cao.

Jin, Northern, Southern dynasties and Three Kingdoms.

After the invasions of northern nomadic peoples at the end of the Western Jin dynasty, the chaos of the Sixteen Kingdoms and the Northern and Southern dynasties ensued. Hebei, firmly in North China and right at the northern frontier, changed hands many times, being controlled at various points in history by the Later Zhao, Former Yan, Former Qin, and Later Yan. The Northern Wei reunified northern China in 440, but split in half in 534, with Hebei coming under the eastern half (first the Eastern Wei; then the Northern Qi), which had its capital at Ye (), near modern Linzhang, Hebei. The Sui dynasty again unified China in 589.

Tang and Five dynasties.

During the Tang dynasty (618–907), the area was formally designated "Hebei" (north of the Yellow River) for the first time. During the earlier part of the Five Dynasties and Ten Kingdoms period, Hebei was fragmented among several regimes, though it was eventually unified by Li Cunxu, who established the Later Tang (923–936). The next dynasty, the Later Jin under Shi Jingtang, posthumously known as Emperor Gaozu of Later Jin, ceded much of modern-day northern Hebei to the Khitan Liao dynasty in the north; this territory, called the Sixteen Prefectures of Yanyun, became a major weakness in the Chinese defense against the Khitans for the next century, since it lay within the Great Wall.

Song, Liao, Jin, and Yuan dynasties.

During the Northern Song dynasty (960–1127), the sixteen ceded prefectures continued to be an area of hot contention between Song China and the Liao dynasty. The Southern Song dynasty that came after abandoned all of North China, including Hebei, to the Jurchen Jin dynasty after the Jingkang Incident in 1127 of the Jin–Song wars.  The Mongol Yuan dynasty divided China into provinces but did not establish Hebei as a province. Rather, the area was directly administrated by the Secretariat () at capital Dadu.

Ming and Qing dynasties.

The Ming dynasty ruled Hebei as "Beizhili" (), meaning "Northern Directly Ruled", because the area contained and was directly ruled by the imperial capital, Beijing; the "Northern" designation was used because there was a southern counterpart covering present-day Jiangsu and Anhui. When the Manchu Qing dynasty came to power in 1644, they abolished the southern counterpart, and Hebei became known as "Zhili", or simply "Directly Ruled". During the Qing dynasty, the northern borders of Zhili extended deep into what is now Inner Mongolia, and overlapped in jurisdiction with the leagues of Inner Mongolia.

Republic of China.

The Qing dynasty collapsed in 1912 and was replaced by the Republic of China. Within a few years, China descended into civil war, with regional warlords vying for power. Since Zhili was so close to Peking (Beijing), the capital, it was the site of frequent wars, including the Zhiwan War, the First Zhifeng War and the Second Zhifeng War. With the success of the Northern Expedition, a successful campaign by the Kuomintang to end the rule of the warlords, the capital was moved from Peking (Beijing) to Nanking (Nanjing). As a result, the name of Zhili was changed to Hebei to reflect the fact that it had a standard provincial administration, and that the capital had been relocated elsewhere.  During the Second World War, Hebei was under the control of the Reorganized National Government of the Republic of China, a puppet state of Imperial Japan.

People's Republic of China.

The founding of the People's Republic of China saw several changes: the region around Chengde, previously part of Rehe Province (historically part of Manchuria), and the region around Zhangjiakou, previously part of Chahar Province (historically part of Inner Mongolia), were merged into Hebei, extending its borders northwards beyond the Great Wall. The capital was also moved from Baoding to the upstart city of Shijiazhuang, and, for a short period, to Tianjin.  On July 28, 1976, Tangshan was struck by a powerful earthquake, the Tangshan earthquake, the deadliest of the 20th century with over 240,000 killed. A series of smaller earthquakes struck the city in the following decade.  Today, Hebei, along with Beijing and Tianjin municipalities which it enclaves, make up the Jing-Jin-Ji megalopolis region. With a population fo 130 million, it is about six times the size as the New York metropolitan area and is one of the largest megalopolis cluster in China. Beijing had also unloaded some of its non-capital functions to the province with the establishment of the Xiong'an New Area, which will further facilitate the integration of the three regions.

Geography.

Most of central and southern Hebei lies within the North China Plain. The western part of Hebei rises into the Taihang Mountains (Taihang Shan), while the Yan Mountains (Yan Shan) run through northern Hebei, beyond which lie the grasslands of Inner Mongolia. The Great Wall of China cuts through northern Hebei from east to west as well, briefly entering the border of Beijing Municipality, and terminates at the seacoast of Shanhaiguan in northeastern Hebei. The highest peak is Mount Xiaowutai () in Yu County in the northwest of the province, with an altitude of.  Hebei borders the Bohai Sea on the east. The Hai He watershed covers most of the province's central and southern parts, and the Luan He watershed covers the northeast. Not counting the numerous reservoirs to be found in Hebei's hills and mountains, the largest lake in Hebei is Baiyangdian, located mostly in Anxin County.  Major cities in Hebei include:

Climate.

Hebei has a continental monsoon climate, with cold, dry winters, and hot, humid summers. Temperatures average in January and in July; the annual precipitation ranges from, concentrated heavily in summer.

Administrative divisions.

Hebei is made up of eleven prefecture-level divisions: all prefecture-level cities:  These eleven prefecture-level divisions are subdivided into 168 county-level divisions (47 districts, 21 county-level cities, 94 counties and 6 autonomous counties). Those are, in turn, divided into 2207 township-level divisions (1 district public office, 937 towns, 979 townships, 55 ethnic townships, and 235 subdistricts). At the end of 2017, the total population of Hebei is 75.2 million.

Politics.

The politics of Hebei is structured in a dual party-government system like all other governing institutions in mainland China.  The Governor of Hebei is the highest-ranking official in the People's Government of Hebei. However, in the province's dual party-government governing system, the Governor has less power than the Hebei Communist Party of China Provincial Committee Secretary (CPC Party Chief).

Economy.

In 2014, Hebei's GDP was 2.942 trillion yuan (US$479 billion), ranked 6th in the PRC. GDP per capita reached 40,124 Renminbi. As of 2011, the primary, secondary, and tertiary sectors of industry contributed 203.46 billion, 877.74 billion, and 537.66 billion RMB respectively. The registered urban unemployment rate was 3.96%.  40% of Hebei's labor force works in the agriculture, forestry and animal husbandry sectors, with the majority of production from these industries going to Beijing and Tianjin. Hebei's main agricultural products are cereal crops including wheat, maize, millet, and sorghum. Cash crops like cotton, peanut, soybeans and sesame are also produced.  Kailuan, with a history of over 100 years, is one of China's first modern coal mines, and remains a major mine with an annual production of over 20 million metric tonnes. Much of the North China Oilfield is found in Hebei, and there are also major iron mines at Handan and Qian'an. Iron, as well as steel, manufacturing are the largest industries in Hebei, and are likely to remain so as these industries consolidate and Hebei continues to grow as a manufacturing and transportation center for the region.  Hebei's industries include textiles, coal, steel, iron, engineering, chemical production, petroleum, power, ceramics and food.

Demographics.

The population is mostly Han Chinese. 55 ethnic minorities are present in Hebei, representing 4.27% of the total population. The largest are Manchu (2.1 million people), Hui people (600000 people) and Mongol (180000 people). Excludes members of the People's Liberation Army in active service. Source: Department of Population, Social, Science and Technology Statistics of the National Bureau of Statistics of China and Department of Economic Development of the State Ethnic Affairs Commission of China, eds. "Tabulation on Nationalities of 2000 Population Census of China". 2 vols. Beijing: Nationalities Publishing House (), 2003. ()  In 2004, the birth rate was 11.98 births per 1,000 people, while the death rate was 6.19 deaths per 1,000 people. In 2000 the sex ratio at birth was 118.46 males to 100 females.

Religion.

The predominant religions in Hebei are Chinese folk religions, Taoist traditions and Chinese Buddhism. According to surveys conducted in 2007 and 2009, 5.52% of the population believe in and are involved in ancestor veneration, while 3.05% of the population identify as Christian, mostly of the Catholic Church. Local worship of deities in the region began to organise into "benevolent churches" as a reaction to Catholicism in the Qing dynasty.  The reports didn't give figures for other types of religion; 90.61% of the population may be either irreligious or involved in worship of nature deities, Buddhism, Confucianism, Taoism and folk religious sects. Zailiism is a folk religious sect that originated in Hebei. There is a presence of Tibetan Buddhist schools in the province.  Hebei has the largest Catholic population in China, with 1 million members according to the local government. and 1.5 million Catholics according to the Catholic Church. The province is considered as the center of Catholicism in China. The town of Donglu, where an apparition of the Virgin Mary was reported to have occurred in 1900, is reportedly "one of the strongholds of the unofficial Catholic Church in China".  A large number of Catholics in Hebei remain loyal to the Pope and reject the authority of the Catholic Patriotic Church. Four of Hebei's underground bishops have been imprisoned in recent years: Bishop Francis An Shuxin of Donglu since 1996; Bishop James Su Zhimin since October 1997; and Bishops Han Dingxiang of Yongnian who died in prison in 2007 and Julius Jia Zhiguo of Zhengding since late 1999. In 2003 there were 350.000 Protestants and 580.000 Muslims according to government statistics. According to a survey, as of 2010 Muslims constitute 0.82% of the population of Hebei.

Culture.

Dialects of Mandarin are spoken over most of the province, and most Mandarin dialects in Hebei are in turn classified as part of the Ji Lu Mandarin subdivision. Regions along the western border with Shanxi, however, have dialects that are distinct enough for linguists to consider them as part of Jin, another subdivision of Chinese, rather than Mandarin. In general, the dialects of Hebei are quite similar to and readily intelligible with the Beijing dialect, which forms the basis for Standard Chinese, the official language of the nation. However, there are also some distinct differences, such as differences in the pronunciation of certain words that derive from entering tone syllables (syllables ending on a plosive) in Middle Chinese.  Traditional forms of Chinese opera in Hebei include Pingju, Hebei Bangzi (also known as Hebei Clapper Opera), and Cangzhou Kuaiban Dagu. Pingju is especially popular: it tends to be colloquial in language and hence easy to understand for audiences. Originating from northeastern Hebei, Pingju has been influenced by other forms of Chinese opera like Beijing opera. Traditionally Pingju makes use of just a "xiaosheng" (young male lead), a "xiaodan" (young female lead), and a "xiaohualian" (young comic character), though it has since diversified with the use of other roles as well.  Quyang County, in central Hebei, is noted for its Dingzhou porcelain, which includes various vessels such as bowls, plates, vases, and cups, as well as figurines. Dingzhou porcelain is usually creamy white, though it is also made in other colours.  Hebei cuisine is typically based on wheat, mutton and beans.

Notable individuals.

Well-known people born in Hebei Province include:

Media.

Hebei is served by the province-wide Hebei Television, abbreviated HEBTV. It also covers parts of Beijing, Tianjin, Shandong, Henan, Shanxi, and Inner Mongolia. Shijiazhuang Radio & Television is a regional TV network that covers the provincial capital. Hebei is also served by three main newspapers, Hebei Daily, Yanzhao Metropolis Daily, and Yanzhao Evening News, all published by the Hebei Daily Newspaper Group.

Transportation.

Because Hebei surrounds Beijing and Tianjin, all the numerous important railway lines radiating out of these two cities pass through Hebei. The Beijing–Guangzhou Railway is one of the most important: it passes through many major cities such as Baoding, Shijiazhuang, Xingtai and Handan on its way south to Henan. Other important railways include the Beijing–Kowloon Railway, Beijing–Shanghai Railway, Beijing-Harbin Railway, Beijing–Chengde Railway, Beijing–Tongliao Railway, Beijing-Baotou Railway and Fengtai–Shacheng Railway. High-speed rail lines crossing the province include the Beijing-Shanghai High-Speed Railway, Beijing-Guangzhou High-Speed Railway and Shijiazhuang–Taiyuan High-Speed Railway. Future high-speed rail lines from Beijing and Tianjin to Northeast China and Northwest China will traverse northern Hebei.  During the Eleventh Five-Year Plan, Beijing and Hebei were collaborating on a new passenger railway. The RMB 82.6 billion network will add 844 kilometers to the system. Current railway systems for Hebei trains are also being upgraded and will soon be able to travel at speeds of between 160 and 200 kilometers per hour.  As of the early 2013, railway schedule systems listed 160 passenger train stations within the province.  The recent expressway boom in China has not left Hebei behind. There are expressways to every prefecture-level city of Hebei, totalling to approximately 2,000 kilometers. The total length of highways within Hebei is around 40,000 kilometers.  There are a number of ports along the Bohai Sea, including Qinhuangdao (the second busiest in China with a capacity of over 100 million tons), Huanghua, and Jingtang. Shijiazhuang's Zhengding Airport is the province's center of air transportation, with domestic and international flights. Parts of Hebei will also be served by the new Beijing Daxing International Airport in Beijing, which is currently under construction and expected to be completed by 2017.

Tourism.

The Ming Great Wall crosses the northern part of Hebei and the eastern end is located on the coast at Shanhaiguan (Shanhai Pass), near Qinhuangdao. Informally known as the "First Pass of The World" (), Shanhaiguan was the place where Ming general Wu Sangui opened the gates to Manchu forces in 1644, beginning nearly 300 years of Manchu rule; Shanhai Pass also marks the psychological entrance / exit of Manchuria, so that for centuries Manchuria was known as "outside the Pass" or "east of the Pass". Beidaihe, located near Shanhaiguan, is a popular beach resort well known as a former meeting place for top governmental officials.  The Chengde Mountain Resort and its outlying temples are a World Heritage Site. Also known as the Rehe Palace, this was the summer resort of the Manchu Qing dynasty emperors. The Chengde Resort was built between 1703 and 1792, and consists of a palace complex, a large park area composed of lakes, pavilions, causeways, bridges, etc., and a number of Tibetan Buddhist and Han Chinese temples in the surrounding area.  There are Qing dynasty imperial tombs at Zunhua (Eastern Qing Tombs) and Yixian (West Qing Tombs). The Eastern Qing Tombs are the resting place of 161 Qing emperors, empresses, and other members of the Qing imperial family, while the West Qing Tombs have 76. These are also part of a World Heritage Site.  The Zhaozhou, or Anji Bridge, built by Li Chun during the Sui dynasty, is the oldest stone arch bridge in China, and one of the most significant examples of pre-modern Chinese civil engineering.  Baoding, the old provincial capital, contains the historical Zhili Governor's Residence.  Xibaipo, a village about from Shijiazhuang, in Pingshan County was the location of the Central Committee of the Communist Party of China and the headquarters of the People's Liberation Army during the decisive stages of the Chinese Civil War between May 26, 1948 and March 23, 1949, at which point they were moved to Beijing. Today, the area houses a memorial site.

Sports.

The 2018 Women's Bandy World Championship was held in Hebei.  Sports teams based in Hebei include:  National Basketball League (China) Hebei Springs Benma  Chinese Football Association

Education.

Under the national Ministry of Education:  Under other national agencies:  Under the provincial government:

Sister regions.

Hebei is twinned with the following states/provinces/prefectures/regions:

Hebei (; alternately Hopeh) is a coastal province of the People's Republic of China, and is part of the North China region. The modern province was established in 1911 as Chihli Province (Zhili Province). Its capital and largest city is Shijiazhuang. Its one-character abbreviation is "" (), named after Ji Province, a Han dynasty province ("zhou") that included what is now southern Hebei. The name "Hebei" literally means "north of the river", referring to its location entirely to the north of the Yellow River.

Hebei (, früher nach Post "Hopeh") ist eine Provinz im Norden der Volksrepublik China. Die Abkürzung für den Provinznamen lautet "jì" (冀), der sich von der Präfektur und dem antiken Staat "Ji" ableitet, in welcher sich die Provinz einmal befand.


Taxonomy and evolution.

The island fox shares the genus "Urocyon" with the mainland gray fox, the species from which it is descended. Its small size is a result of insular dwarfism, a form of allopatric speciation. Because the island fox is geographically isolated, it has no immunity to parasites and diseases brought in from the mainland and is especially vulnerable to those that the domestic dog may carry. In addition, predation by the golden eagle and human activities devastated fox numbers on several of the Channel Islands in the 1990s. Four island fox subspecies were federally protected as an endangered species in 2004, and efforts to rebuild fox populations and restore the ecosystems of the Channel Islands are being undertaken. Radio collars are being attached to foxes in an effort to track and locate the young foxes. To date these efforts have been largely successful.  There are six subspecies of the island fox, each of which is native to a specific Channel Island, and which evolved there independently of the others. The subspecies are:  Foxes from each island are capable of interbreeding, but have genetic and phenotypic distinctions that make them unique; for example, the subspecies have differing numbers of tail vertebrae.  The small size of the island fox is an adaptation to the limited resources available in the island environment. The foxes are believed to have "rafted" to the northern islands between 10,400 and 16,000 years ago. Initially, fox populations were located on the three northern islands, which were likely easier to access during the last ice age — when lowered sea levels united four of the northernmost islands into a single mega-island (Santa Rosae) and the distance between the islands and the mainland was reduced — it is likely that Native Americans brought the foxes to the southern islands of the archipelago, perhaps as pets, or hunting dogs. Other names for the island fox include "coast fox", "short-tailed fox", "island gray fox", "Channel Islands fox", "Channel Islands gray fox", "California Channel Islands fox" and "insular gray fox".

Description.

The island fox is significantly smaller than the gray fox and is the smallest fox in North America, averaging slightly smaller than the swift and kit foxes. Typically, the head-and-body length is, shoulder height, and the tail is long, which is notably shorter than the tail of the gray fox. This is due to the fact that the island fox generally has two fewer tail vertebrae than the gray fox. The island fox weighs between. The male is always larger than the female. The largest of the subspecies occurs on Santa Catalina Island and the smallest on Santa Cruz Island.  The island fox has gray fur on its head, a ruddy red coloring on its sides, white fur on its belly, throat and the lower half of its face, and a black stripe on the dorsal surface of its tail. In general the coat is darker and duller hued than that of the gray fox. The island fox molts once a year between August and November. Before the first molt pups are woolly and have a generally darker coat than adult foxes. A brown phase, with the grey and black fur of the body replaced by a sandy brown and a deeper brown, may occur in the San Clemente Island and San Nicolas Island populations. It is unclear if this is a true color phase, a change that occurs with age, or possibly a change that occurs because of interactions with Opuntia cactus spines that become embedded in the pelt.

Reproduction.

The island fox typically forms monogamous breeding pairs, which are frequently seen together beginning in January and through the breeding season, from late February to early March. The gestation period is 50–63 days. The female island fox gives birth in a den, a typical litter having one to five pups, with an average of two or three. Pups are born in the spring and emerge from the den in early summer; the mother lactates for 7–9 weeks. Sexual maturity is reached at 10 months, and the females usually breed within the first year. Island foxes live for 4–6 years in the wild and for up to 8 years in captivity.

Ecology and behavior.

Its preferred habitat is complex layer vegetation with a high density of woody, perennially fruiting shrubs. The fox lives in all of the island biomes including temperate forest, temperate grassland and chaparral, with no island supporting more than 1,000 foxes. The island fox eats fruits, insects, birds, eggs, crabs, lizards, and small mammals, including deer mice. The fox tends to move around by itself, rather than in packs. It is generally nocturnal, although with peaks of activity at dawn and dusk. Activity also fluctuates with the season: It is more active during the day in summer than it is in winter.  The island fox is not intimidated by humans, although at first may show aggression. It is quite easy to tame and is generally docile. The island fox communicates using auditory, olfactory and visual signals. A dominant fox uses vocalizations, staring, and ear flattening to cause another fox to submit. Signs of dominance and submission are visual, such as facial expression and body posture. Its main vocalizations are barking and growling. The island fox marks territory with urine and feces.

Conservation status and Federal Protection.

In March 2004, four subspecies of the island fox were classified as a federally protected endangered species: The Santa Cruz island fox, Santa Rosa island fox, San Miguel island fox and the Santa Catalina island fox. As of 2013, the IUCN lists the entire species as near threatened, an improvement from its previous status of "critically endangered". A decline in island fox populations was identified in the 1990s. On San Miguel Island, the decline began in 1994, with the population falling from 450 adults to 15 by 1999. Similar population declines were discovered on Santa Cruz Island. On the Santa Cruz Island the population decreased from 2,000 adults in 1994 to less than 135 in 2000, and on Santa Rosa Island where foxes may have numbered more than 1,500 in 1994, but were reduced to 14 animals by 2000. In 2004, there were 38 San Miguel island foxes, all in captivity; 46 foxes in captivity on Santa Rosa Island and 7 in the wild (golden eagle predation prevented the release of captive foxes into the wild); Santa Cruz Island had 25 captive foxes and a stable wild population of around 100 foxes.  Golden eagle predation, discovered when foxes were radio-collared and monitored, proved to be the cause of the high mortality rates. The golden eagle was an uncommon visitor to the Channel Islands before the 1990s according to data gathered by Dr. Lyndal Laughrin of the University of California Santa Cruz Island Reserve, and the first golden eagle nest was recorded on Santa Cruz Island in 1999. Biologists propose that the eagle may have been attracted to the islands in the 1960s after the decline of the bald eagle. The golden eagle replaced the bald eagle and began to feed on feral pigs following the decimation of the local bald eagle population due to DDT exposure in the 1950s — the bald eagle would have deterred the golden eagle from settling on the islands while it subsisted on fish.  The feral pigs on Santa Rosa were exterminated by the National Park Service in the early 1990s, which removed one of the golden eagle's food sources. The golden eagle then began to prey on the island fox population. Feral pigs on Santa Cruz Island and deer and elk on Santa Rosa Island were introduced almost 70 years prior to island fox decline, therefore, the golden eagle most likely did not seek these animals as alternative prey. This has occurred most likely as a result of a process known as "apparent competition": In this process, a predator, like the golden eagle, feeds on at least two prey, for example, the island fox and feral pigs. One prey item is adapted to high predation pressure and supports the predator population (i.e. pigs), whereas the other prey item (i.e. the island fox) is poorly adapted to predation and declines as a consequence of the predation pressure. It has also been proposed that the complete removal of golden eagles may be the only action that could save three subspecies of the island fox from extinction. However, the pigs on Santa Cruz Island were killed by the Nature Conservancy under the idea that they drew the eagles to the foxes.  Introduced diseases or parasites can devastate island fox populations. Because the island fox is isolated, it has no immunity to parasites and diseases brought in from the mainland and are especially vulnerable to those the domestic dog may carry. A canine distemper outbreak in 1998 killed approximately 90% of Santa Catalina Island's foxes, reducing the population from 1,300 to 103 in 2000. A vaccination program has been initiated to protect Catalina Island foxes from canine distemper. After several years of carefully trapping the foxes and vaccinating them against distemper and rabies, their population has reached 1,717 in 2015, surpassing the pre-disease population of about 1,300. Scientists believe the distemper virus was introduced by a pet dog or a raccoon from the mainland that hitched a ride on a boat or a barge. To eliminate the risk of disease, pets are not permitted in Channel Islands National Park.  Diminished food supply and general degradation of the habitat due to introduced mammal species, including feral cats, pigs, sheep, goats, and American bison, the latter having been introduced to Catalina Island in the 1920s by a Hollywood film crew shooting a Western, also has had a negative effect on fox populations.  The foxes threaten a population of the severely endangered San Clemente Island loggerhead shrike in residence on San Clemente Island. The island fox population has been negatively affected by trapping and removal or euthanasia of foxes by the United States Navy. Since 2000, the Navy has employed different management strategies: Trapping and holding foxes during the shrike breeding season, the installation of an electric fence system around shrike habitats, and the use of shock collar systems. With the gradual recovery of the shrike population on San Clemente Island, the Navy no longer controls the foxes.  The populations of Santa Cruz island foxes, San Miguel island foxes, and Santa Rosa island foxes have dramatically rebounded from lows in 2000 of 70 for the Santa Cruz foxes and 15 each on San Miguel and Santa Rosa Islands. The Catalina Island Conservancy runs a captive breeding program on Catalina Island. On September 14, 2012, the US Fish and Wildlife Service released a draft recovery plan for the San Miguel island fox, Santa Rosa island fox, Santa Cruz island fox, and the Santa Catalina island fox. By 2012, the Catalina Island Conservancy determined that there were 1,500 Santa Catalina island foxes and the population was stable. As of 2015, there were 520 native foxes on San Miguel and 874 on Santa Rosa, according to the group Friends of the Island Fox. The number of foxes on Santa Cruz Island had risen to 1,750. The US Fish and Wildlife Service recommended delisting Santa Cruz, San Miguel and Santa Rosa island foxes in a major success of the Endangered Species Act. However, they are recommending that the Santa Catalina island be reclassified from endangered to threatened, because of the threat of diseases on this heavily visited island.  Two other subspecies on San Nicolas and San Clemente are not endangered. There were 263 foxes on San Nicolas and 1,230 on San Clemente.  Because the Channel Islands are almost entirely owned and controlled by either the Catalina Island Conservancy, The Nature Conservancy, or the federal government, the fox has a chance to receive the protection it needs, including constant supervision by interested officials without the ongoing threat of human encroachment on its habitat.  The fox did not persist on Anacapa Island because it has no reliable source of fresh water; Santa Barbara Island is too small to support the food needs of a viable fox population.  Rene Vellanoweth, an archaeologist, believes that inbreeding depression can be managed by mixing the different island fox subspecies populations much as the indigenous peoples did, by moving them from island to island, creating a higher genetic diversity and assisting them in recovery.

The island fox ("Urocyon littoralis") is a small fox that is native to six of the eight Channel Islands of California. There are six subspecies, each unique to the island it lives on, reflecting its evolutionary history. Island foxes are generally docile, show little fear of humans, and are easily tamed. Island foxes played an important role in the spiritual lives of native Channel Islanders. Island foxes have been likely semi-domesticated as pets, used as pelts, or for other functions, like pest control.

Der Insel-Graufuchs oder Kalifornische Insel-Graufuchs ("Urocyon littoralis") ist eine Art der Graufüchse. Er lebt nur auf sechs der acht Kanalinseln vor der Küste Kaliforniens und ist damit eine endemische Art.


Types.

A person's prosthesis should be designed and assembled according to the person's appearance and functional needs. For instance, a person may need a transradial prosthesis, but need to choose between an aesthetic functional device, a myoelectric device, a body-powered device, or an activity specific device. The person's future goals and economical capabilities may help him or her choose between one or more devices.  Craniofacial prostheses include intra-oral and extra-oral prostheses. Extra-oral prostheses are further divided into hemifacial, auricular (ear), nasal, orbital and ocular. Intra-oral prostheses include dental prostheses such as dentures, obturators, and dental implants.  Prostheses of the neck include larynx substitutes, trachea and upper esophageal replacements,  Somato prostheses of the torso include breast prostheses which may be either single or bilateral, full breast devices or nipple prostheses.  Penile prostheses are used to treat erectile dysfunction, correct penile deformity, perform phalloplasty and metoidioplasty procedures in biological men, and to build a new penis in female-to-male gender reassignment surgeries.

Limb prostheses.

Limb prostheses include both upper- and lower-extremity prostheses.  Upper-extremity prostheses are used at varying levels of amputation: forequarter, shoulder disarticulation, transhumeral prosthesis, elbow disarticulation, transradial prosthesis, wrist disarticulation, full hand, partial hand, finger, partial finger. A transradial prosthesis is an artificial limb that replaces an arm missing below the elbow.  Upper limb prostheses can be categorized in three main categories: Passive devices, Body Powered devices, Externally Powered (myoelectric) devices. Passive devices can either be passive hands, mainly used for cosmetic purpose, or passive tools, mainly used for specific activities (e.g. leisure or vocational). An extensive overview and classification of passive devices can be found in a literature review by Maat "et.al." A passive device can be static, meaning the device has no movable parts, or it can be adjustable, meaning its configuration can be adjusted (e.g. adjustable hand opening). Despite the absence of active grasping, passive devices are very useful in bimanual tasks that require fixation or support of an object, or for gesticulation in social interaction. According to scientific data a third of the upper limb amputees worldwide use a passive prosthetic hand. Body Powered or cable operated limbs work by attaching a harness and cable around the opposite shoulder of the damaged arm. The third category of prosthetic devices available are myoelectric arms. These work by sensing, via electrodes, when the muscles in the upper arm move, causing an artificial hand to open or close. In the prosthetics industry, a trans-radial prosthetic arm is often referred to as a "BE" or below elbow prosthesis.  Lower-extremity prostheses provide replacements at varying levels of amputation. These include hip disarticulation, transfemoral prosthesis, knee disarticulation, transtibial prosthesis, Syme's amputation, foot, partial foot, and toe.

History.

Prosthetics originates from the ancient Near East circa 3000 BC, with the earliest evidence of prosthetics appearing in ancient Egypt and Iran. The earliest recorded mention of eye prosthetics is from the Egyptian story of the Eye of Horus dates circa 3000 BC, which involves the left eye of Horus being plucked out and then restored by Thoth. Circa 3000-2800 BC, the earliest archaeological evidence of prosthetics is found in ancient Iran, where an eye prosthetic is found buried with a woman in Shahr-i Shōkhta. It was likely made of bitumen paste that was covered with a thin layer of gold. The Egyptians were also early pioneers of foot prosthetics, as shown by the wooden toe found on a body from the New Kingdom circa 1000 BC. Another early recorded mention is found in South Asia circa 1200 BC, involving the warrior queen Vishpala in the Rigveda. Roman bronze crowns have also been found, but their use could have been more aesthetic than medical.  An early mention of a prosthetic comes from the Greek historian Herodotus, who tells the story of Hegesistratus, a Greek diviner who cut off his own foot to escape his Spartan captors and replaced it with a wooden one.

Wood and metal prosthetics.

Pliny the Elder also recorded the tale of a Roman general, Marcus Sergius, whose right hand was cut off while campaigning and had an iron hand made to hold his shield so that he could return to battle. A famous and quite refined historical prosthetic arm was that of Götz von Berlichingen, made at the beginning of the 16th century. The first confirmed use of a prosthetic device, however, is from 950–710 BC. In 2000, research pathologists discovered a mummy from this period buried in the Egyptian necropolis near ancient Thebes that possessed an artificial big toe. This toe, consisting of wood and leather, exhibited evidence of use. When reproduced by bio-mechanical engineers in 2011, researchers discovered that this ancient prosthetic enabled its wearer to walk both barefoot and in Egyptian style

Technology progress before the 20th century.

An Italian surgeon recorded the existence of an amputee who had an arm that allowed him to remove his hat, open his purse, and sign his name. Improvement in amputation surgery and prosthetic design came at the hands of Ambroise Paré. Among his inventions was an above-knee device that was a kneeling

Lower extremity modern history.

After the Second World War a team at the University of California, Berkeley including James Foort and C.W. Radcliff helped to develop the quadrilateral socket by developing a jig fitting system for amputations above the knee. Socket technology for lower extremity limbs saw a further revolution during the 1980s when John Sabolich C.P.O., invented the Contoured Adducted Trochanteric-Controlled Alignment Method (CATCAM) socket, later to evolve into the Sabolich Socket. He followed the direction of Ivan Long and Ossur Christensen as they developed alternatives to the quadrilateral socket, which in turn followed the open ended plug socket, created from wood. The advancement was due to the difference in the socket to patient contact model. Prior to this, sockets were made in the shape of a square shape with no specialized containment for muscular tissue. New designs thus help to lock in the bony anatomy, locking it into place and distributing the weight evenly over the existing limb as well as the musculature of the patient. Ischial containment

Upper extremity modern history.

In 2005, DARPA

Patient procedure.

A prosthesis is a functional replacement for an amputated or congenitally malformed or missing limb. Prosthetists are responsible for the prescription, design, and management of a prosthetic device.  In most cases, the prosthetist begins

Current technology and manufacturing.

Over the years, there have been advancements in artificial limbs. New plastics and other materials, such as carbon fiber, have allowed artificial limbs to be stronger and lighter, limiting the amount of extra energy necessary to operate the limb. This is especially important for trans-femoral amputees. Additional materials have allowed artificial limbs to look much more realistic, which is important to trans-radial and transhumeral amputees because they are more likely to have the artificial limb exposed. In addition to new materials, the use of electronics has become very common in artificial limbs. Myoelectric limbs, which control the limbs by converting muscle movements to electrical signals, have become much more common than cable operated limbs. Myoelectric signals are picked up by electrodes, the signal gets integrated and once it exceeds a certain threshold, the prosthetic limb control signal is triggered which is why inherently, all myoelectric controls lag. Conversely, cable control is immediate and physical, and through that offers a certain degree of direct force feedback that myoelectric control does not. Computers are also used extensively in the manufacturing of limbs. Computer Aided Design and Computer Aided Manufacturing are often used to assist in the design and manufacture of artificial limbs.  Most modern artificial limbs are attached to the residual limb (stump) of the amputee by belts and cuffs or by suction. The residual limb either directly fits into a socket on the prosthetic, or—more commonly today—a liner is used that then is fixed to the socket either by vacuum (suction sockets) or a pin lock. Liners are soft and by that, they can create a far better suction fit than hard sockets. Silicone liners can be obtained in standard sizes, mostly with a circular (round) cross section, but for any other residual limb shape, custom liners can be made. The socket is custom made to fit the residual limb and to distribute the forces of the artificial limb across the area of the residual limb (rather than just one small spot), which helps reduce wear on the residual limb. The custom socket is created by taking a plaster cast of the residual limb or, more commonly today, of the liner worn over the residual limb, and then making a mold from the plaster cast. Newer methods include laser-guided measuring which can be input directly to a computer allowing for a more sophisticated design.  One problem with the residual limb and socket attachment is that a bad fit will reduce the area of contact between the residual limb and socket or liner, and increase pockets between residual limb skin and socket or liner. Pressure then is higher, which can be painful. Air pockets can allow sweat to accumulate that can soften the skin. Ultimately, this is a frequent cause for itchy skin rashes. Over time, this can lead to breakdown of the skin.  Artificial limbs are typically manufactured using the following steps:

Attachment to the body.

Most prostheses can be attached to the exterior of the body, in a non-permanent way. Some others however can be attached in a permanent way. One such example are exoprostheses (see below).

Direct bone attachment and osseointegration.

Osseointegration is a method of attaching the artificial limb to the body. This method is also sometimes referred to as exoprosthesis (attaching an artificial limb to the bone), or endo-exoprosthesis.  The stump and socket method can cause significant pain in the amputee, which is why the direct bone attachment has been explored extensively. The method works by inserting a titanium bolt

Cosmesis.

Cosmetic prosthesis has long been used to disguise injuries and disfigurements. With advances in modern technology, cosmesis, the creation of lifelike limbs made from silicone or PVC has been made possible. Such prosthetics, including artificial hands, can now be designed to simulate the appearance of real hands, complete with freckles, veins, hair, fingerprints and even tattoos.

Cognition.

Unlike neuromotor prostheses, neurocognitive prostheses would sense or modulate neural function in order to physically reconstitute or augment cognitive processes such as executive function, attention, language, and memory. No neurocognitive prostheses are currently available but the development of implantable neurocognitive brain-computer interfaces has been proposed to help treat conditions

Prosthetic enhancement.

In addition to the standard artificial limb for everyday use, many amputees or congenital patients have special limbs and devices to aid in the participation of sports and recreational activities.  Within science fiction, and, more recently, within the scientific community, there has been consideration given to using advanced prostheses to replace healthy body parts with artificial mechanisms and systems to improve function. The morality and desirability of such technologies are being debated by transhumanists, other ethicists, and others in general. Body parts such as legs, arms, hands, feet, and others can be replaced.  The first experiment with a healthy individual appears to have been that by the British scientist Kevin Warwick. In 2002, an implant was interfaced directly into Warwick's nervous system. The electrode array, which contained around a hundred electrodes, was placed in the median nerve. The signals produced were detailed enough that a robot arm was able to mimic the actions of Warwick's own arm and provide a form of touch feedback again via the implant.  The DEKA company of Dean Kamen developed the "Luke arm", an advanced nerve-controlled prosthetic. Clinical trials began in 2008, with FDA approval in 2014 and commercial manufacturing by the Universal Instruments Corporation expected in 2017. The price offered at retail by Mobius Bionics is expected to be around $100,000.  Further research in April 2019, there have been improvements towards prosthetic function and comfort of 3D-printed personalized wearable systems. Instead of manual integration after printing, integrating electronic sensors at the intersection between a prosthetic and the wearer's tissue can gather information such as pressure across wearer's tissue, that can help improve further iteration of the these types of prosthetic.

Oscar Pistorius.

In early 2008, Oscar Pistorius, the "Blade Runner" of South Africa, was briefly ruled ineligible to compete in the 2008 Summer Olympics because his transtibial prosthesis limbs were said to give him an unfair advantage over runners who had ankles. One researcher found that his limbs used twenty-five percent less energy than those of an able-bodied runner moving at the same speed. This ruling was overturned on appeal, with the appellate court stating that the overall set of advantages and disadvantages of Pistorius' limbs

Design considerations.

There are multiple factors to consider when designing a transtibial prosthesis. Manufacturers must make choices about their priorities regarding these factors.

Performance.

Nonetheless, there are certain elements of socket and foot mechanics

Other.

The buyer is also

Cost and source freedom.

High-cost.

In the USA a typical prosthetic limb costs anywhere between $15,000 and $90,000, depending on the type of limb desired by the patient. With medical insurance, a patient will typically pay 10%–50% of the total cost of a prosthetic limb, while the insurance company will cover the rest of the cost. The percent that the patient pays varies on the type of insurance plan, as well as the limb requested by the patient. In the United Kingdom, much of Europe, Australia and New Zealand the entire cost of prosthetic limbs is met by state funding or statutory insurance. For example, in Australia prostheses are fully funded by state schemes in the case of amputation due to disease, and by workers compensation or traffic injury insurance in the case of most traumatic amputations. The National Disability Insurance Scheme, which is being rolled out nationally between 2017 and 2020 also pays for

Low-cost.

Low-cost above-knee prostheses often provide only basic structural support with limited function. This function is often achieved with crude, non-articulating, unstable, or manually locking knee joints. A limited number of organizations, such as the International Committee of the Red Cross (ICRC), create devices for developing countries. Their device which is manufactured by CR Equipments is a single-axis, manually operated locking polymer prosthetic knee joint.  Table. List of knee joint technologies based on the literature review.  A plan for a low-cost artificial leg, designed by Sébastien Dubois, was featured at the 2007 International Design Exhibition and award show in Copenhagen, Denmark, where it won the. It would be able to create an energy-return prosthetic leg

Open-source robotic prothesis.

There is currently an open-design Prosthetics forum known as the "Open Prosthetics Project". The group employs collaborators and volunteers to advance Prosthetics technology while attempting to lower the costs of these necessary devices. Open Bionics is a company that is developing open-source robotic prosthetic hands. It uses 3D printing to manufacture

Low-cost prosthetics for children.

In the USA an estimate was found of 32,500 children (<21 years) that suffer from major paediatric amputation, with 5,525 new cases each year, of which 3,315 congenital.  Carr et al. (1998) investigated amputations caused by landmines for Afghanistan, Bosnia and Herzegovina, Cambodia and Mozambique among children (<14 years), showing estimates of respectively 4.7, 0.19, 1.11 and 0.67 per 1000 children. Mohan (1986) indicated in India a total of 424,000 amputees (23,500 annually), of which 10.3% had an onset of disability below the age of 14, amounting to a total of about 43,700 limb deficient children in India alone.  Few low-cost solutions have been created specially for children. Examples of low-cost prosthetic devices include:

Pole and crutch.

This hand-held pole with leather support band or platform for the limb is one of the simplest and cheapest solutions

Bamboo, PVC or plaster limbs.

This also fairly simple solution comprises a plaster socket with a bamboo or PVC pipe at the bottom, optionally attached to a prosthetic

Adjustable bicycle limb.

This solution is built using a bicycle seat post up side down

Sathi Limb.

It is an endoskeletal modular lower limb from India,

Monolimb.

Monolimbs are non-modular prostheses and thus require more experienced prosthetist for correct fitting,

Cultural and social theory perspectives.

A number of theorists have explored the meaning and implications of prosthetic extension of the body. Elizabeth Grosz writes, "Creatures use tools, ornaments, and appliances to augment their bodily capacities. Are their bodies lacking something, which they need to replace with artificial or substitute organs?...Or conversely, should prostheses be understood, in terms of aesthetic reorganization and proliferation, as the consequence of an inventiveness that functions beyond and perhaps in defiance of pragmatic need?" Elaine Scarry argues that every artifact recreates and extends the body. Chairs supplement the skeleton, tools append the hands, clothing augments the skin. In Scarry's thinking, "furniture and houses are neither more nor less interior to the human body than the food it absorbs, nor are they fundamentally different from such sophisticated prosthetics as artificial lungs, eyes and kidneys. The consumption of manufactured things turns the body inside out, opening it up "to" and "as" the culture of objects." Mark Wigley, a professor of architecture, continues this line of thinking about how architecture supplements our natural capabilities, and argues that "a blurring of identity is produced by all prostheses." Some of this work relies on Freud's earlier characterization of man's relation to objects as one of extension.

In medicine, a prosthesis (plural: prostheses; from Ancient Greek "prosthesis", "addition, application, attachment") or prosthetic implant is an artificial device that replaces a missing body part, which may be lost through trauma, disease, or a condition present at birth (congenital disorder). Prostheses are intended to restore the normal functions of the missing body part. Amputee rehabilitation is primarily coordinated by a physiatrist as part of an inter-disciplinary team consisting of physiatrists, prosthetists, nurses, physical therapists, and occupational therapists. Prostheses can be created by hand or with computer-aided design (CAD), a software interface that helps creators design and analyze the creation with computer-generated 2-D and 3-D graphics as well as analysis and optimization tools.

Eine Prothese (von, ‚Anfügung‘, aus und ), auch Körperersatzstück, bezeichnet ein Gliedmaßen, Organe oder Organteile ersetzendes, künstlich geschaffenes, funktionell ähnliches Produkt.