 After schooling at Vienna and Kraków, he graduated from the Jan Kazimierz University in Lviv in 1928 and worked for a while in the Department of Comparative Anatomy.He received a PhD in 1932, working under Kazimierz Kwietniewski on the anatomy of the urinary tract and associated glands of mice.At the end of 1938 he moved to the Jagiellonian University as assistant professor of zoology.