 One explanation is their higher incidence of gene and whole genome duplication and retention of those additional genes, due in part to their development of a large collection of defensive secondary metabolites.The apparent disconnect between the number of genes in a species and its biological complexity was dubbed the G-value paradox.While the C-value paradox unraveled with the discovery of massive sequences of noncoding DNA, resolution of the G-value paradox appears to rest on differences in genome productivity.