 in Entomology from Purdue University in 1994 and a Ph.D. in Entomology from University of Wisconsin-Madison in 1999.During his PhD program he spent time, as a visiting student, at both the University of Chicago in the Department of Ecology and Evolution and at CSIRO at Black Mountain in Canberra, Australia.He completed his post-doctoral training at the Max Planck Institute for Chemical Ecology in Jena, Germany, in 2000.