The great leonopteryx (Toruk in Navi, meaning last shadow) is the apex airborne predator native to Pandora. It is scientifically known as Leonopteryx rex, meaning "king lion-wing". The fierce beauty and nobility of the leonopteryx gave the species a place central to Navi lore and culture. The leonopteryx is scarlet with black stripes and a midnight blue crest on top of the head and on the lower jaw. It is celebrated in dance and song; elaborate totems symbolize both the fear and respect accorded to the creature. The leonopteryx is considered crucial to the Navi sense of destiny and interconnectedness. Prior to the events of the film, it had only been tamed five times, and Neytiri tells Jake that the riders (Toruk Makto) brought peace among the Pandoran tribes. This makes Jake the sixth Toruk Makto. He manages to capture one by jumping on it from above from his banshee but releases it after the battle with the humans has ended. The great leonopteryx and the banshee were designed with bright colors. Page based the colors on Earths birds, poison dart frogs, and monarch butterflies, though he altered the patterns so that their inspirations would not be so conspicuous to moviegoers. The skull and crest shapes appear to be derived from pterosaurs from the genus Tapejara.