While there is no enrichment of S between trophic levels, the stable isotope can be useful in distinguishing benthic vs. pelagic producers and marsh vs. phytoplankton producers. Similar to C, it can also help distinguish between different phytoplankton as the key primary producers in food webs. The differences between seawater sulfates and sulfides (c. 21‰ vs -10‰) aid scientists in the discriminations. Sulfur tends to be more plentiful in less aerobic areas, such as benthic systems and marsh plants, than the pelagic and more aerobic systems. Thus, in the benthic systems, there are smaller δS values.