In 1962, Harvard professor Edward O. Wilson published one of the first concrete studies constructing the groundwork for the notion of trail pheromones. Claiming an odor trail is deposited by the sting apparatus of the hymenopteran Solenopsis saevissima which results in a pathway from the colony to a food source, this study encouraged further investigation of how this chemical is laid, how it affects communication between species within and outside of its own, the evolution of the semiochemical, etc.