Three proteinaceous iron–sulfur reaction centers are found in PSI. Labeled F, F, and F, they serve as electron relays. F and F are bound to protein subunits of the PSI complex and F is tied to the PSI complex. Various experiments have shown some disparity between theories of iron–sulfur cofactor orientation and operation order. In one model, F passes an electron to F, which passes it on to F to reach the ferredoxin.