In dead Sphagnum, sphagnan a polysaccharide with D-lyxo-5-hexosulouronic acid is a major remaining substance. It makes the bog very acidic, so that bacteria cannot grow. Not only that, the plant ensures there is no available nitrogen. Holocellulose also absorbs any digestive enzymes around. Together this leads to major accumulation of peat under sphagnum bogs.